In 1996 we detailed the organization and activation of vertebrate behavior and included molecular epigenetics. The model was extended to invertebrates and led to details of nutrient chemical-dependent pheromone-controlled social and sexual behavior in species from microbes to man.
The honeybee model organism provides the best example of the epigenetic tweaking of immense gene networks in superorganisms that solve problems through the exchange and the selective cancellation and modification of signals. However, this primate research makes it clearer how an environmental drive evolved from that of food ingestion in unicellular organisms to that of socialization in insects and in mammals.
In primates and other mammals, food odors and pheromones cause changes in hormones such as LH, which has developmental affects on behavior in nutrient-dependent, reproductively fit individuals across species that signal fitness via pheromones. Ingested estrogens->pheromones->behavior, get it?